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rary human Tx/91 and N. grill 99 Cal/99 H1N1 viruses, which require an exogenous protease source for their multicycle replication and plaque formation (Table 1), the 1918 virus and a recombinant in-fluenza virus bearing the 1918 HA and NA seg-ments only (1918 HA/NA:Tx/91) formed visible plaques without the addition of trypsin (Table 1). surya 99